Please e-mail for reprints.

Refereed journal articles:


Plowright RK, Becker DJ, Crowley DE, Washburne AD, Huang T, Han BA. 2019. Nipah virus outbreak in Kerala, India: Identifying reservoirs to prioritize surveillance. PLoS Neglected Tropical Diseases, 13(6): e0007393.

Schmidt JP, Maher S, Drake JM, Huang T, Farrell MJ, Han BA. 2019. Ecological indicators of mammal exposure to Ebolavirus. Philosophical Transactions of the Royal Society B. 374:20180337.

Berger KM, Wood JLN, Jenkins B, Olsen J, Morse SS, Gresham L, Root JJ, Rush M, Pigott D, Winkleman T, Gillespie TR, Nuzzo J, Han BA, Olinger P, Karesh WB, Mills JN, Annelli JF, Barnabei J, Lucey D, Hayman DTS. 2019. Policy and science for global health security: shaping the course of international health. Special issue: One Health and Zoonoses, Topical Medicine and Infectious Diseases 4(2): 60.

Han BA, Majumdar S, Calmon FD, Horesh R, Kumar A, Perer A, von Marschall EB, Wei D, Mojsilovic A, Varshney K. 2019. Confronting data sparsity to identify potential sources of Zika virus infection among primates. Epidemics, 27:59.

Han BA, Ostfeld RS. 2019. Topic modeling of major research themes in disease ecology of mammals. Journal of Mammalogy, 100:1008.

Downs C, Schoenle L, Han BA, Harrison J, Martin M. 2019. The scaling of host competence. Trends in Parasitology, 35:182.

Stephens PR, Altizer S, Gittleman JL, Moan E, Han BA, Pappalardo P. 2019. Parasite sharing in wild ungulates and their predators: effects of phylogeny, range overlap, and trophic links. Journal of Animal Ecology, 19:1159.

Dallas TA, Han BA, Nunn CL, Park AW, Stephens PR, Drake JM. 2019. Host traits associated with species roles in parasite sharing networks. Oikos 28:23.

Almeida RM, Han BA, Reisinger AJ, Kagemann C, Rosi EJ. 2018. High mortality in aquatic predators of mosquito larvae caused by exposure to insect repellent. Biology Letters 14, 20180526.

Dallas TA, Aguirre AA, Budischak S, Carlson C, Ezenwa V, Han BA, Huang S, Stephens PR. 2018. Gauging support for macroecological patterns in helminth parasites. Global Ecology and Biogeography 27, 1437–1447.

Walker JW, Han BA, Ott IM and Drake JM. 2018. Transmissibility of emerging viral zoonoses. PLoS ONE. 13, e0206926.

Strona G, Carstens CJ, Beck PSA, Han BA. 2018. The intrinsic vulnerability of networks to epidemics. Ecological Modelling 383:91–97. doi:10.1016/j.ecolmodel.2018.05.013

Yang LH, Han BA. 2018. Data-driven predictions and novel hypotheses about zoonotic tick vectors from the genus Ixodes. BMC Ecology, 18:1-7.

      [PDF] [DATA]

Blaustein AR, Urbina J, Snyder PW, Reynolds E, Dang T, Hoverman JT, Han BA, Olson DH, Searle C, Hambalek NH. 2018. The effects of emerging infectious diseases on amphibians: a review of experimental studies. Diversity 10, 81.


LaDeau SL, Han BA, Rosi-Marshall EJ, Weathers KC. 2017. The next decade of big data in ecosystem science. Ecosystems. 20:274–83. 

Evans, M., Dallas, T.A., Han, B.A., Murdock, C.C., Drake, J.M. Data-driven identification of potential Zika virus vectors. 2017.  eLife. DOI:


Schmidt, J.P., Park, A.W., Kramer, A., Han, B.A., Alexander, L.A., Draker, J.D. 2017 . Spatiotemporal fluctuations and triggers of Ebola virus spillover. Emerging Infectious Diseases. DOI: 10.3201/eid2303.160101. [Link to Visualization].


Han, B.A., Yang, L. 2016Predicting novel tick vectors of zoonotic disease. Proceedings of the 33rd International Conference on Machine Learning.  


Pigott, D.M., Millear, A., Earl, L., Morozoff, C., Han, B.A., Shearer, F.M., Weiss, D.J., Brady, O.J., Kraemer, M.U.G., Moyes, C.L., Bhatt, S., Gething, P.W., Golding, N., Hay, S.I. 2016. Updates to the zoonotic niche map of Ebola virus disease in Africa. eLife, 5:e16412


Han, B.A., Schmidt, J.P., Hayman, D.T.S., Alexander, L.A., Bowden, S.A., Drake, J.M. 2016. Undiscovered bat hosts of filoviruses. PLoS Neglected Tropical Diseases, 10(7): e0004815.

[Press: Digital Trends, Scientific Computing, ScienceDaily, Infection Control Today, Foreign Affairs New Zealand]


Stephens, P.R., Altizer, S.M., Smith, K.F., Aguirre, A., Brown, J.H., Budischak, S., Byers, J.E., Critchlow, R., Davies, J.T., Drake, J.M., Ezenwa, V., Farrell, M., Gittleman, J.L., Han, B.A., Huang, S., Hutchinson, R.A., Johnson, P.T.J., Nunn, C.L., Onstad, D., Park, A.P., Poulin, R., Vazquez-Prokopec, G.M., Papparlardo, P.,Schmidt, J.P. 2016. The macroecology of infectious diseases: a new perspective on global-scale drivers of pathogen distributions and impacts. Ecology Letters, 19:1159.



Han, B.A., Kramer, A., Drake, J.M. 2016. Global patterns of zoonotic disease in mammals. Trends in Parasitology, 32:565-577

[Data] [PDF]

[Press: BBC World Report, The washington PostThe Scientist, The Science Explorer, Conservation magazine, SINC, El País, TakePart, Cell Press Podcast, WAMC Northeast public radio, Popular Science, Health Medicine Network, Tech Times, MSN Noticias, FiercePharma, Eurek Alert, Le Scienze]


Han, B.A., Drake, J.M. 2016. Future directions in analytics for infectious disease intelligence. EMBO Reports, 2016:e201642534. DOI: 10.15252/embr.201642534



LaDeau, S., Han, B.A. 2016. The emergence of disease ecology. Japanese Journal of Zoo and Wildlife Management. 21(3)



Han, B.A., Schmidt, J.P., Bowden, S.E., Drake, J.M. 2015Rodent reservoirs of future zoonotic diseases. Proceedings of the National Academy of Science, 112(22):7039-7044. doi:10.1073/pnas.1501598112

[Data] [Peer Reviews] [PDF]

[Press: ScienceNature, The EconomistIEEE Spectrum, The ScientistFusionHealioHomeland Security News WirePopular Science, Infection Control TodayLaboratory EquipmentNews MedicalOnlineAthens, Poughkeepsie JournalRT, Ars Technica, National Geographic, PHYS ]


Han, B.A., Park, A.W., Jolles, A.E., Altizer, S. 2015Infectious diseases transmission and behavioural allometry in wild mammals. Journal of Animal Ecology, 84(3):637-646. doi:10.1111/1365-2656.12336

[Data] [Peer Reviews] [PDF]


Han, B.A., Kerby, J.L., Searle, C.L., Storfer, A., Blaustein, A.R. 2015Host species composition influences infection severity among amphibians in the absence of spillover transmission. Ecology and Evolution, 5(7):1432-1439. doi:10.1002/ece3.1385

[Data] [Peer Reviews] [PDF]


Kats, L.B., Bucciarelli, G., Schlais, D.E., Blaustein, A.R., and Han, B.A. 2012Ultraviolet radiation influences perch selection by a neotropical poison-dart frog. PLoS ONE, 7:e51364. doi:10.1371/journal.pone.0051364



Han, B.A., Searle, C.L., and Blaustein, A.R. 2011The effects of an infectious fungal pathogen, Batrachochytrium dendrobatidis, on amphibian predator-prey interactions. PLoS ONE, 6(2): e16675.  doi:10.1371/journal.pone.0016675



Altizer, S., Bartel, R., and Han, B.A. 2011. Animal migrations and infectious disease risk. Science, 331:296-302.


[Press: Faculty of 1000, LiveScience, ScienceDaily, Scientific American, TIME Magazine, US News & World Report]


Blaustein, A.R., Han, B.A., Relyea, R., Johnson, P.T.J., Buck, J., Gervasi, S. and Kats, L.B. 2011. The complexity of amphibian population declines: understanding the role of cofactors in driving amphibian losses. Annals of the New York Academy of Sciences, The Year in Ecology and Conservation Biology (Eds. Ostfeld, R.S. and Schlesinger, W.H.), 1223:108-119. doi: 10.1111/j.1749-6632.2010.05909.x



Bancroft, B.A., Han, B.A., Searle, C.L., Biga, L.M., Olson, D.H., Kats, L.B., Lawler, J.J., and Blaustein, A.R. 2011. Species-level correlates of susceptibility to the pathogenic amphibian fungus Batrachochytrium dendrobatidis in the United States. Biodiversity and Conservation, 20:1911-1920. doi: 10.1007/s10531-011-0066-4



Romansic, J.R., Johnson, P.T.J., Searle, C.L., Johnson, J.E., Tunstall, T., Han, B.A., Rohr, J.R., and Blaustein, A.R. 2011. Individual and combined effects of multiple pathogens on Pacific treefrogs. Oecologia, 166(4):1029-1041. doi: 10.1007/s00442-011-1932-1



Searle, C.L., Belden, L.K., Bancroft, B.A., Han, B.A., Biga, L.F., and Blaustein, A.R. 2010. Experimental examination of the effects of ultraviolet-B radiation in combination with other stressors in frog larvae. Oecologia, 162:237-245.



Han, B.A., Bradley, P.W., and Blaustein, A.R. 2008. Ancient behaviors of larval amphibians in response to an emerging fungal pathogen, Batrachochytrium dendrobatidis. Behavioral Ecology and Sociobiology, 63:241-250.



Lampo, M., Sánchez, D., Nicolás, A., Márquez, M., Nava-González, F., Garcia, C.Z., Rinaldi, M., Rodríguez-Contreras, A., León, Fabiola, Han, B.A., Chacón-Ortiz, A. 2008. Batrachochytrium dendrobatidis in Venezuela. Herpetological Review, 39:449-454.



Sánchez, D.A., Chacón-Ortiz, A., León, R., Han, B.A., and Lampo, M. 2008. Widespread occurrence of an emerging pathogen in amphibian communities of the Venezuelan Andes. Biological Conservation, 141:2898-2905.



Han, B.A., Kats, L.B., Pommerening, R.C., Ferrer, R.P., Murry-Ewers, M. and Blaustein A.R. 2007. Behavioral avoidance of ultraviolet-B radiation by two species of neotropical poison-dart frogs. Biotropica, 39:433-435.


[Press: Conservation Magazine, Grist, Mongabay]


Lampo, M., Barrio-Amoros, C.L., and Han, B.A. 2006. Batrachochytrium dendrobatidis infection in the recently rediscovered Atelopus mucubajiensis (Anura, Bufonidae) in the Venezuelan Andes. EcoHealth, 3:299-302.



Johnson, P.T.J., Preu, E. R., Sutherland, D. R., Romansic, J., Han, B.A., and Blaustein, A.R. 2006. Adding infection to injury: Synergistic effects of predation and parasitism on salamander limb malformations. Ecology, 87:2227–2235.



Blaustein, A. R., Romansic, J. M., Scheessele, E. A., Han, B.A., Pessier, A.P., and Longcore, J.E. 2005. Interspecific variation in susceptibility of frog tadpoles to the pathogenic fungus Batrachochytrium dendrobatidis. Conservation Biology, 19:1460-1468.



Blaustein, A.R., Han, B., Fasy, B., Romansic, J., Scheessele, E.A., Anthony, R.G., Marco, A., Chivers, D.P., Belden, L.K., Kiesecker, J.M., Garcia, T.S., Lizana, M. and Kats, L.B. 2004. Variable breeding phenology affects the exposure of amphibian embryos to ultraviolet radiation and Optical characteristics of natural waters protect amphibians from UV-B in the U.S. Pacific Northwest: Comment. Ecology, 85:1747-1754. 


Other publications:

Pandit P, Han BA. Rise of the Machines in Disease Ecology: The Arising and Established Researcher. Bulletin of the Ecological Society of America, January 2020.

Han BAThe Algorithm That's Hunting Ebola. Invited feature article, IEEE Spectrum Magazine, 24 September 2015.


Han BA and Altizer S. 2013Invited chapterConservation and Diseases in The Encyclopedia of Biodiversity (2nd edition). Levin, S. (Ed.) Academic Press. doi:10.1016/B978-0-12-384719-5.00273-2



Han BA, Rushmore J, Fritzsche A, Satterfield D, and Winternitz J. 2012Preempting pandemics. Science, 337:647-648. (Book Review: The Viral Storm by Nathan Wolfe). doi:10.1126/science.1225462